Full References and Abstracts of Mycological Articles

Soop K., 1986: En jämförelse mellan två spindelskivlingar inom Phlegmacium sektion Amarescentes Moser — Jordstjärnan 7 (1): 4-5.

Summary:
A comparison between two Cortinarii in subg. Phlegmacium sectio Amarescentes Moser. Cortinarius subtortus Fr. is compared with finds of C. amurceus Fr. from northern Pinus forest, and it is shown that the two taxa can be maintained at a specific level.


Soop K., 1986: Några observationer av spindelskivlingar inom Sericeocybe grupp Anomali — Jordstjärnan 7 (2): 5-10.

Notes on Cortinarii in subg. Sericeocybe group Anomali.


Soop K., 1987: Notes et observations sur les Cortinaires de Suède — Documents Mycologiques 68: 45-64.
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Les espèces suivantes sont traitées :


Soop K., 1988: Notes et Observations sur les Cortinaires de Belgique — Documents Mycologiques 73: 23-32.
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Les espèces suivantes sont traitées :


Soop K., 1987: Cortinarius armeniacus Fr. och närstående arter — Jordstjärnan 8(2): 28-34.
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Summary:
Cortinarius armeniacus and related species. A small group around Cortinarius armeniacus is reviewed. The taxa belong to the subgenus Telamonia, they have a brownish cap and whitish, non-radicating stipe, and grow in spruce forests. Taxa with distinctly coloured flesh or stipe are explicitly excluded. Despite the bewildering variety within even such a circumscribed group, most of the finds in Sweden turn out to be either C. armeniacus Fr. ss latu, or C. dilutus Fr. Several collections of a red-brown to date-brown form of C. armeniacus are reported, and it is argued that this taxon is in reality identical to C. triformis Fr. ss Mos., Marchand. The form is in fact mentioned by Fries [1830] as a variety of C. armeniacus under the name Ag. badius Schum.

As regards C. melleo-pallens (Fr.) Lge, the author has several collections that might be attributed to this taxon, but it is argued that there are very few, if any, characters that can be unambiguously used to distinguish it from armeniacus. A review of the common literature shows that authors tend to assign more or less identical characters to the two. Although it is possible that Lange has found a pale variety of C. triformis Fr. ss Fr nec auct, as witnessed by his icon, it is certainly not the taxon referenced by most authors. Fries' own triformis (including his form melleopallens) is probably a rare species in Sweden.

The following species are discussed:

Soop K., 1989: Notes et observations sur les Cortinaires (Suite) — Documents Mycologiques 77: 77-91.
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Les espèces suivantes sont traitées :


Soop K., 1988: Cortinarius cremeolaniger Orton found in Sweden — Agarica 18: 92-95.
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Soop K., 1989: Intressanta spindlingar från Uppsalatrakten — Jordstjärnan 10(1): 15-18.
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Summary:
Interesting Cortinars from the Uppsala area. During a foray in the vicinity of Upsala, the author found a number of interesting Cortinarii: (1) what might be the southernmost find of C. canabarba Mos. in the Nordic countries; (2) several collections of C. (Phlegm.) russus Fr., characterised by its telamonioid aspect and nauseating taste, in calcareous Picea forest; and (3) C. (Leprocybe) cf. subnotatus Fr., a darkening, olivaceus species, in a Corylus wood. The two latter species are described and discussed. In particular, species (3) appears related to C. uraceus Fr. and to C. viridipes Mos., and is probably identical to C. uraceus ss Melot (1986), though the spores appear slightly shorter.

Soop K., 1990: The group Cortinarius paragaudis Fries in Sweden — Agarica 10/11 (19/20) : 98-107.
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Summary:
The paper attempts to clarify the specific variety in the group containing C. paragaudis Fr. and C. hæmatochelis Bull.:Fr. An analysis of twelve collections from Central Sweden is presented, and it is shown that the material can be split roughly into two groups, separated mainly by spore size. Finally, a new species, resembling a C. paragaudis with a white veil, is described. The following species are described:


Soop K., 1990: Un nouveau Cortinaire de la bétulaie subalpine — Documents Mycologiques 80: 39-42.

Summary:
A new Cortinarius from subalpine Betula forests. A new Cortinarius (Telamonia) with a reddening veil is described. The species appears related to C. spilomeus Fr. var. depauperatus Lge, but differs in an initially greyish veil with a faint olivaceous hue. Lange's taxon is promoted to specific rank and moved to Telamonia, where the two species form a natural group, to which one should possibly add C. heterosporus Bres.

  • Cortinarius depauperatus (Lge)Soop stat. nov.
  • Cortinarius heterocyclus Soop sp. nov. (with colour photo)


Soop K., 1990: Observationer av några intressanta Cortinarius-arter — Agarica 10/11 (19/20) : 30-43.
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Summary:
Notes on some interesting species of Cortinarius. The paper describes a number of interesting Cortinarii from Central Sweden. Several rare or critical species are described in detail with references to illustrative material. In some cases (C. impennis, lucorum, salor ssp. transiens), a revised taxonomy is proposed. The following species are discussed:


Soop K., 1993: On Cortinarius in boreal pine forests — Agarica 12(21): 101-116 .
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Summary:
The paper reports on observations of a number of interesting Cortinarii in boreal forests with special emphasis on the Pinus habitat. Similar taxa or forms that occur with Pinus and Picea are compared. Two new species are described and a few taxa are re-interpreted. The following species are described:


Soop K., 1993: Notes et observations sur les Cortinaires de Suède (suite) — Documents Mycologiques 90: 9-15.
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Summary:
A number of Cortinarii described in a series of earlier articles (Soop [1987-89]) are revisited and reinterpreted along with several new, interesting finds.


Soop K., 1994: En ny titt på spindlingar inom gruppen Smör-&-Bröd — Jordstjärnan 15(1): 12-19.
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Summary:
A new look at Cortinars in the Bread-&-Butter group. A difficult group of Cortinarii may be designated "bread-&-butter" due to their deceptively commonplace look and ubiquitous presence. A restricted set of such taxa is examined: they are all middle-sized Telamoniæ with a white veil, brownish cap and pale stipe, growing with conifers. New varieties of C. tortuosus (Fr.:Fr.)Fr. and C. biformis Fr. are described. Taxa illustrated by colour photographs:


Soop K., 1995: Smör-&-bröd spindlingar, del II — Jordstjärnan 16(2): 30-36.
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Summary:
Bread-&-Butter Cortinars, part II. This paper continues the theme of difficult Cortinarii that may be designated "bread-&-butter" due to their deceptively commonplace look and ubiquitous presence. The most frequently encountered Telamoniæ with a white veil, brownish cap, and a pale stipe seem to be adequately presented in modern literature. Not all of these trivially looking species are common, however. In this instalment a few uncommon ones are described and illustrated in colour:


Soop K., 1997: Spindlingar med röd slöja — Jordstjärnan 18(1): 26-35.
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Summary:
Cortinarii with a red veil. A key is given to Scandinavian species of Cortinarius with a reddish (red or brown-red to vinaceous) veil, and four uncommon and interesting species are described and illustrated in colour. The key, which is partly based on M. Moser's well-known key of red-veiled Cortinarii (Moser 1965), concentrates on the subgenus Telamonia. The following species are described and illustrated by colour photos:


Soop K., 1998: Cortinarius på Undersidan — Jordstjärnan 19(1): 25-37.
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Summary:
Cortinarius Down Under. Observations of taxa in Cortinariaceae from a three-week field trip to the South Island of New Zealand are reported. After a comparison of the Northern and Southern Hemispheres in terms of biotopes, mycorrhiza, and foraying conditions in general, a number of taxonomic problems at the infra- and suprageneric level of Cortinarius are noted.
The following species are illustrated in colour:


Soop K., 1998: Fårskär vid Upplandskusten -- en märklig granskogslokal för spindlingar — Jordstjärnan 19(2): 38-43.
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Summary:
Fårskär on the Uppland Coast — a remarkable coniferous locality for Cortinarius. A list of Cortinarius finds is presented from an extremely productive, calcareous Picea locality in Northern Uppland near the Baltic shoreline. Many of the finds (checkmarked in the list) are on the red list of endangered species in Sweden and/or Norway.

Two of the finds are described and illustrated in colour photographs: C. bovinus Fr. is interpreted according to M. Moser (as opposed to ss Lange, a taxon which is considered identical to C. bulbosus Fr.). The second species, C. cæsiostramineus, differs from a similar taxon, also found in the locality, by a (sometimes strong) blue coloration on the stipe, a more fibrillose cap, lack of a bitter taste (except possibly as an after-taste), a fruity smell, and slightly shorter spores. The latter taxon, identified as C. amarescens Mos., is sometimes regarded as a synonym, but should be separated at least by varietal rank. Finally a beautifully orange-coloured species akin to C. callisteus Fr. is depicted, differing from the former by colour, by (almost) white gills when undeveloped, lack of odor, and markedly stronger fluorescence in UV light. It might be recognised as a form of C. tophaceoides Mos. ined., which however has the typical stearine smell of callisteus, or possibly as Flammula abrupta Fr. ss Melot. The following species are illustrated by colour photos:


Soop K., 1998: Notes et observations sur les champignons cortinarioïdes de Nouvelle-Zélande — Documents Mycologiques 112: 13-26.

Summary:
Notes and observations on cortinarioid fungi of New Zealand. The problems relating to the supra- and infrageneric taxonomy of the genus Cortinarius are examined with regard to the contribution of the Southern Hemisphere, without, however, discerning a definite solution. Six new cortinarioid species from New Zealand are presented. The following species are illustrated by colour photos:


Soop K., 2000: Images des champignons cortinarioïdes de Nouvelle-Zélande — Journal des JEC 2: 63-69.

Texte accompagnant une projection d'images d'espèces cortinarioïdes et autres, provenant des forêts spontanées de l'Île du Sud, présentée à la réunion des J.E.C. en 1999.

Summary:
Pictures of cortinarioid fungi from New Zealand. This is the text part of a slide show of cortinarioid and other species from the native forests on the South Island of New Zealand, presented at the JEC meeting in 1999.


Soop K., 2002: Una revisione del complesso Cortinarius suberi — Boll. Micol. Bresadola 44(3): 7-12.
English translation p. 12-14.
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Summary:
A review of the Cortinarius suberi complex. The macro- and microscopic characters of two Cortinarius taxa in section Malachii, C. suberi and C. brunneogriseus are re-examined, and it is concluded that they should be separated only at the varietal level. Detailed descriptions and colour illustrations are provided of:


Soop K., 2001: Les cortinaires séricéocyboïdes des forêts de résineux des pays nordiques — Journal des JEC 3: 3-9.
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Résumé
Les Sericeocybe (s. l.) forment un élément important dans les forêts de conifères en Scandinavie. Cet article donne une clé pour le groupe et en décrit trois taxons critiques et rares avec des illustrations en couleur.

Summary
The sericeocyboid Cortinarii in coniferous forests of the Nordic countries. Sericeocybe (s. l.) forms an important element in Scandinavian coniferous forests. The paper gives a key for the group and describes three critical and rare taxa with colour illustrations.

Les taxons suivants sont traités et illustrés :


Soop K., 2001: Contribution à l'étude de la mycoflore cortinarioïde de Nouvelle-Zélande — Bull. Soc. Mycol. France 117(2): 91-132.

Résumé
Vingt-huit nouveaux taxons sont décrits des forêts spontanées de la Nouvelle-Zélande, dont 23 Cortinarius, 1 Thaxterogaster, 2 Pholiota, 1 Gymnopilus et 1 Hebeloma.

Summary
A contribution to the study of the cortinarioid mycoflora of New Zealand. Twenty-eight new taxa are described from the native forests of New Zealand, of which 23 Cortinarius, 1 Thaxterogaster, 2 Pholiota, 1 Gymnopilus and 1 Hebeloma.

Les taxons suivants sont décrits ; certains illustrés par des photos en couleur :

  • Cortinarius anauensis Soop
  • Cortinarius aurantioferreus Soop (avec photo)
  • Cortinarius austrocyanites Soop
  • Cortinarius carbonellus Soop
  • Cortinarius caryotis Soop
  • Cortinarius collybianus Soop
  • Cortinarius cremeolina Soop
  • Cortinarius cupreonatus Soop
  • Cortinarius dysodes Soop
  • Cortinarius exlugubris Soop
  • Cortinarius lubricanescens Soop
  • Cortinarius minoscaurus Soop
  • Cortinarius naphthalinus Soop
  • Cortinarius papaver Soop (avec photo)
  • Cortinarius perelegans Soop (avec photo)
  • Cortinarius periclymenus Soop
  • Cortinarius persicanus Soop (avec photo)
  • Cortinarius picoides Soop
  • Cortinarius rattinus Soop
  • Cortinarius rotundisporus ssp. nothofagi Soop (avec photo)
  • Cortinarius saturniorum Soop
  • Cortinarius ursus Soop (avec photo)
  • Cortinarius viscilætus Soop
  • Gymnopilus rubrocastaneus Soop
  • Hebeloma mediorufum Soop
  • Pholiota chrysmoides Soop
  • Pholiota multicingulata var. hanmerensis Soop
  • Thaxterogaster viola Soop

Soop K., 2002: Contribution à l'étude de la mycoflore cortinarioïde de Nouvelle-Zélande, II — Bull. Soc. Mycol. France 118(3): 173-194.

Résumé
Treize nouveaux taxons, dont une nouvelle section, de Cortinarius sont décrits des forêts spontanées de la Nouvelle-Zélande.

Summary
A contribution to the study of the cortinarioid mycoflora of New Zealand, II. Thirteen new Cortinarius taxa, including one new section, are described from the native forests of New Zealand.

Les taxons suivants sont décrits ; certains illustrés par des photos en couleur :

  • Cortinarius sectio Xenosmatæ Soop
  • Cortinarius chalybæus Soop (avec photo)
  • Cortinarius cretax Soop
  • Cortinarius ignellus Soop
  • Cortinarius incensus Soop
  • Cortinarius iringa Soop (avec photo)
  • Cortinarius kaimanawa Soop
  • Cortinarius luteinus Soop
  • Cortinarius subgemmeus Soop
  • Cortinarius suecicolor Soop
  • Cortinarius tigrellus Soop
  • Cortinarius veronicæ v. dilutus Soop
  • Cortinarius xenosma Soop (avec photo)

Soop K., 2005: A contribution to the study of the cortinarioid mycoflora of New Zealand, III

Summary
Ten new species and one new variety of Cortinarius are described from the native forests of New Zealand. The following taxa are described:

  • Cortinarius alboaggregatus Soop
  • Cortinarius castoreus Soop
  • Cortinarius elaiochrous var. leontis Soop
  • Cortinarius eutactus Soop
  • Cortinarius ionomataius Soop
  • Cortinarius lamproxanthus Soop
  • Cortinarius laquellus Soop
  • Cortinarius paraonui Soop
  • Cortinarius paraxanthus Soop
  • Cortinarius singularis Soop
  • Cortinarius thaumastus Soop

B. Gasparini & K. Soop, 2008: Contribution to the knowledge of Cortinarius [Agaricales, Cortinariaceae] of Tasmania (Australia) and New Zealand. — Australasian Mycologist 27(3): 173-203.
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Summary
The authors have been studying the mycological flora of New Zealand and Tasmania over many years with the aim of discovering phylogenetic links between Cortinarius occurring in the Southern and Northern Hemispheres. Based on morphological and molecular phylogenetic studies, which will be published elsewhere, we describe here ten new species of Cortinarius from Tasmania and eleven from New Zealand. Further we rename two taxa and describe one additional species. The following new taxa are described and illustrated in colour:

The following taxa are renamed: The following taxon is discussed in detail:

K. Soop, 2010(2013): A contribution to the study of the cortinarioid mycoflora of New Zealand, IV — Bresadoliana,1(2) : 17–34.
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Summary
Ten new species and one new variety of Cortinarius are described from the native forests of New Zealand. The following taxa are described:


K. Soop, 2013: A contribution to the study of the cortinarioid mycoflora of New Zealand, V — Australasian Mycologist 31: 1-9.
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Summary
Six new species and one new variety are described from New Zealand. The following taxa are described:


K. Soop, 2014: A contribution to the study of the cortinarioid mycoflora of New Zealand, VI — NZ Journ. Bot. 52(3): 328-342.
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Summary
Ten new species are described from New Zealand and are provided with keys. The following taxa are described:


Karl Soop, Michael Wallace and Bálint Dima, 2018: New Cortinarius (Agaricales) species described from New Zealand — NZ Journ. Bot. 56(2): 163-182.
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Summary
As part of an ongoing effort to investigate the diversity of the genus Cortinarius in New Zealand, 12 new species are described. Moreover, the diversity of Cortinarius section Limonii and subgenus Callistei are reviewed in the light of recent findings in the South Pacific, and a key is provided. It is shown that both infrageneric taxa are bihemispherical, whereas the core population of the former is actually located in New Zealand. A phylogram, covering the taxa discussed, is presented to illustrate their genetic relations. The following taxa are described:


K. Soop, 2016: A contribution to the study of the cortinarioid mycoflora of New Zealand, VII — NZ Journ. Bot. 54(3): 344-365.
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Summary
Fourteen new species and two new sections are described from New Zealand Also, a new form of C. lubricanescens is described, and the Patagonian taxon C. rapaceus var. luridus is raised to species rank under a replacement name. The following taxa are described:


Soop K, Dima B, Szarkàndi JG, Cooper J, Papp T, Vàgvölgyi Cs, Nagy LG, 2016: Psathyloma, a new genus in Hymenogastraceae described from New Zealand — Mycologia 108(2): 397Ð404.
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Summary
A new genus, Psathyloma E. Horak ex Soop, J.A. Cooper & Dima, and two new species are described from New Zealand:


K. Soop, 2014: Quelques Telamonia intéressants du nord de l'Europe, III — Journal JEC 16: 3-9.
English translation p. 219-221.
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Résumé
Nous décrivons deux espèces rares et intéressantes du groupe des Telamonia séricéocyboïdes de la Suède centrale : Cortinarius franchii sp. nov., et C. suillus, espèce friesienne quelque peu énigmatique, qui a reçu diverses interprétations. Ces espèces sont illustrées par des photographies en couleur.

Summary
Some interesting Telamoniæ from Northern Europe, III. Two interesting and rare species in the group of sericeocyboid Telamoniæ are described from Central Sweden: Cortinarius franchii sp. nov., and the Friesian C. suillus, a rather enigmatic taxon that has received various interpretations. These species are illustrated with colour photos.


Brandrud TE, Dima B, Schmidt-Stohn G, Bellù F, Frøslev T, Oertel B, Saar G, Soop K, 2014: Cortinarius subgenus Phlegmacium section Multiformes in Europe — Journal JEC 16: 162-199.
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Summary
The taxonomy and phylogeny of sect. Multiformes (Cortinarius, subgen. Phlegmacium) are presented. The present circumscription of the section is supported by molecular data and morphological features. The taxonomy is complex and many taxa are quite new and little known. Two new species are described. Altogether ten European species are recognized: C. multiformis, C. talimultiformis, C. frondosomultiformis sp. nov., C. rufoallutus, C. talus, C. melleicarneus, C. caesiolamellatus, C. caesiophylloides, C. pallidirimosus and C. armenicorius sp. nov. The ten recognized species are (i) more or less well-supported phylogenetically, but (ii) with a fairly low morphological differentiation. The species differs by over 1% in ITS DNA from sister taxa, and each species pair show a morphological differentiation, though in most cases with overlapping characters. For instance, most species have overlapping variation in size and shape of the spores, and less than half of the species have formerly been recognized in morphological studies.

Résumé
Cortinarius sous-genre Phlegmacium section Multiformes en Europe. On présente la taxinomie et la phylogénie de la section Multiformes (Cortinarius, sous-genre Phlegmacium). La délimitation actuelle de la section est étayée par des données moléculaires et des caractères morphologiques. La taxinomie est compliquée et beaucoup d'espèces sont tout à fait nouvelles et peu connues. On reconnait en tout 10 espèces européennes : C. multiformis, C. talimultiformis, C. frondosomultiformis sp. nov., C. rufoallutus, C. talus, C. melleocarneus, C. caesiolamellatus, C. caesiophylloides, C. pallidirimosus et C. armenicorius sp. nov. Ces 10 espèces sont (i) plus ou moins bien identifiées au plan phylogénétique, toutefois (ii) assez peu différenciées morphologiquement. Les espèces diffèrent des espèces sœurs à raison de plus de 1% quant à leur ITS d'ADN, et chaque paire d'espèces présente certes une différenciation morphologique, mais avec des chevauchements. C'est ainsi que les dimensions et la forme des spores de la plupart des espèces sont très semblables, et moins de la moitié des espèces présentées ici étaient autrefois distinguées dans les études morphologiques.


Saar G, Brandrud TE, Dima B, Schmidt-Stohn G, Bellù F, Frøslev T, Oertel B, Soop K, 2015: Cortinarius sous-genre Phlegmacium section Purpurascentes en Europe — Journal JEC 17:135-148.
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Résumé
Les données de génétique moléculaire afférentes à de nombreuses récoltes du genre Cortinarius, sous-genre Phlegmacium, section Purpurascentes autorisent maintenant une nouvelle façon de considérer ce groupe. Un phylogramme donne une vue d'ensemble des espèces qu'on peut délimiter aujourd'hui. Pour caractériser la section, on peut aussi recourir à des caractères morphologiques (par exemple les changements de couleur de la chair à l'air) et à des réactions macrochimiques (par exemple de la chair au lugol). En l'état actuel des connaissances, la section Purpurascentes englobe 5 espèces, dont on présente des photos, des descriptions et une clé de détermination.


Redeuilh G. & Soop K., 2006: Nomenclature et taxinomie des genres affines à Fistulinella (Boletaceæ) et Fistulinella lutea sp. nov. de Nouvelle-Zélande — Bull. Soc. Mycol. France 122(4): 291-304.

Résumé
Essai de clarification du complexe constitué par les genres Fistulinella, Ixechinus, Prophyrellus, Mucilopilus et Tylopilus. Description d'une nouvelle espèce récoltée en Nouvelle-Zélande, Fistulinella lutea sp. nov.

Summary
Nomenclature and taxonomy of the genera related to Fistulinella (Boletaceæ) and Fistulinella lutea sp. nov. from New Zealand. It is attempted to clarify the complex of genera consisting of Fistulinella, Ixechinus, Prophyrellus, Mucilopilus and Tylopilus. A new species, found in New Zealand, Fistulinella lutea sp. nov. is described.


Soop K., 2003: Nya rön om släktskapen inom Cortinariaceæ (spindlingar och närstående) — Jordstjärnan 24(1): 28-43.
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Summary
New data on affinities within Cortinariaceæ. In recent times a flurry of papers have reported on molecular-biological studies of taxa in this family. After a review of the concepts and methods involved, the results are summarised for a number of groups (clades) with emphasis on Cortinarius ss. lato, and their impact on current taxonomy is assessed. It is recognised that any attempt at this diverse and complex group must include extensive material from the Southern Hemisphere. The practical consequences for European taxonomy are likely to be limited, however.


Soop K., 2002: Une étude du mycota cortinarioïde des pinèdes sablonneuses dans les pays nordiques — Journal des JEC 4: 3-16.
Deutsche Übersetzung S. 17-27.
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Résumé
Malgré leur importance comme un élément dominant, les cortinaires qui poussent par préférence dans les plantations sablonneuses de Pinus sylvestris en Fennoscandie boréale n'ont pas été l'objet fréquent d'études. Le présent article discute plusieurs groupes et espèces caractéristiques du milieu et en décrit une nouvelle espèce, les illustrant avec des photos en couleur. Une clé est fournie à la fin.

Summary
A study of the cortinarioid mycota in sandy pine forests of the Nordic Countries. Despite their importance as a dominant element, Cortinarius growing in sandy plantations of Pinus sylvestris in boreal Fennoscandia have not been subject to many studies. This paper discusses several characteristic groups and species from this environment, describes one new species, and illustrates them with colour photos. A key is given at the end.

Les taxons suivants sont traités et illustrés :


Soop K., 2003: Cortinarius latus Fries, une espèce critique et intéressante — Journal des JEC 5: 8-11.
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Résumé
Plusieurs récoltes d'un Phlegmacium, apparemment très rare, sont décrites et illustrées par des photos en couleur. Après un examen critique du nom Cortinarius latus, nous concluons qu'il s'applique à notre taxon.

Summary
Cortinarius latus Fries, a critical and interesting species. Several finds of an apparently very rare Phlegmacium are described and illustrated by colour photos. After critically examining the name Cortinarius latus, we conclude that it applies to our taxon.


Soop K., 2003: Première récolte de Cortinarius subtigrinus Reumaux en Suède — Journal des JEC 5: 3-5.
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Résumé
Ce Telamonia rare a été récolté dans une station connue pour sa mycoflore fascinante. Il est caractérisé par les ceintures noir brunâtre sur le stipe, évoquant C. fuscoperonatus Mos.

Summary
First find of Cortinarius subtigrinus Reumaux in Sweden. This rare Telamonia was found in a location known for its fascinating mycoflora. It is caracterised by its blackish-brown bands on the stipe, making it resemble C. fuscoperonatus Mos.


Soop K., 2005: Quelques cortinaires appartenant aux Sericeocybe et groupes voisins rencontrés sous les feuillus. — Journal des JEC 7: 9-17.
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Résumé
Nous décrivons ici trois espèces intéressantes de Cortinarius rencontrées en Suède : C. diabolicus, C. arvalis, et C. cremeolaniger. Elles ont en commun un port robuste, un revêtement sec et pâle et l'habitat sous feuillus. Ces espèces sont comparées entre elles et avec une quatrième, C. ortonii ; elles sont illustrés par des photographies en couleurs.

Summary
A few sericeocyboid Cortinarius found in frondose woods. We describe three interesting Cortinarius found in Sweden: C. diabolicus, C. arvalis, and C. cremeolaniger. They share several caracters: a robust stature, a dry and pale surface, and growing with frondose trees. The species are compared among themselves as well as with a fourth species, C. ortonii. These cortinars are illustrated by colour photos.


Soop K., 2009: Quelques Telamonia intéressants du nord de l'Europe — Journal des JEC 11: 5-9.
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Résumé
Nous décrivons ici deux espèces intéressantes de Cortinarius rencontrées en Suède : C. biveloides Henry des bétulaies, et C. panellus Soop n. sp. des pinèdes sablonneuses. Ces cortinaires sont illustrés par des photos en couleur.

Summary
A few Telamonia from Northern Europe. We here describe two species of Cortinarius found in Sweden: C. biveloides Henry in birch woods, and C. panellus Soop n. sp. in sandy pine forests. These cortinars are illustrated by colour photos.


Soop K., 2010: Quelques Telamonia intéressants du nord de l'Europe, II — Journal des JEC 12: 3-10.
English translation p. 82-86.
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Résumé
Une clé des Cortinarius subg. Telamonia à lames rougeâtres et de taille moyenne, rencontrés dans les pessières boréales est fournie. Deux nouvelles espèces : Cortinarius mattiæ et C. privignatus sont proposées, et une rare espèce, C. redactus Britz., est décrite. Ces taxons sont illustrés par des photos en couleur.

Summary
Some interesting Telamoniæ from Northern Europe, II. A key to red-gilled, medium-sized Cortinarius subg. Telamonia, encountered in boreal Picea forests, is provided. Two new species: Cortinarius mattiæ and C. privignatus are proposed, and a rare species, C. redactus Britz., is described. These taxa are illustrated with colour photos.


Soop K. & Gasparini B., 2011: Europe et Pacifique Sud: Une comparaison de deux flores de Cortinarius — Journal des JEC 13: 34-51.
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Résumé
Nous passons en revue la diversité de Cortinarius et le recouvrement des taxons, aux différents rangs taxonomiques et dans deux zones diamétralement opposées du globe, à la lumière de plusieurs études moléculaires récentes ainsi que de nos propres observations sur le terrain. Les résultats sont illustrés par des photos d’espèces choisies du Pacifique Sud.

Summary
Europe and the South Pacific: A comparison of two Cortinarius floras. We review the diversity and commonality of Cortinarius, at different taxonomical ranks and in two diametrically opposite areas of the globe, in the light of several recent molecular studies as well as of our own field observations. The results are illustrated by photos of species selected from the South Pacific.


Lindström, H. & Soop K., 1999: Quelques petits Telamonia calciphiles — Journal des JEC 1: 40-60.
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A series of small calciphilous Telamoniæ. Seven Cortinarius from subgenus Telamonia, of which five are new, are presented, principally from two Scandinavian habitats, namely calcareous, sandy pine forests and frondose pastures. We describe two taxa from the latter biotope which seem to form mychorrhiza with Helianthemum. The following taxa are described and illustrated by colour photos:


Jacobsson S. & Soop K., 2000: A review of Cortinarii with boletoid spores — Journal des JEC 2:3-12.
Traduction française p. 13-18.
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Summary:
Taxa with lean, narrowly fusoid spores in the genus Cortinarius are reviewed and the discussion focusses on the interpretation of two names: C. heterosporus Bres. and C. aureifolius Peck. Type material and own collections have been investigated, including SEM studies and DNA sequencing. The name C. heterosporus is lectotypified. Colour photographs of both species are provided.


Soop K. & Knutsson T., 2005: Un nouveau cortinaire de la section Calochroi. — Journal des JEC 7: 51-55.
English translation p. 56-58.
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Résumé
Cortinarius ionodactylus sp. nov. est décrit des terrains calcaires secs à végétation peu exigeante en eau et à essence forestière dominante de Corylus du sud-est de la Suède. La nouvelle espèce est illustrée par des photos en couleur.

Summary
A new Cortinarius in Section Calochroi. Cortinarius ionodactylus sp. nov. is described from xerophytic, mosaic woodland dominated by Corylus on calcareous soil in South-Eastern Sweden. The new species is illustrated by colour photos.


Soop K., 1990-97: Ovanliga Cortinarius-arter — en bildserie, del I-IX — Jordstjärnan:

Uncommon Cortinarii — a Picture Series. The following Cortinarius species are described, commented, and depicted in colour photos:

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Part 2, 11(1): 65-67 ladda ner pdf (2000 KB) Part 3, 11(3): 14-15 ladda ner pdf (650 KB) Part 4, 12(2): 16-18 ladda ner pdf (1500 KB) Part 5, 12(3): 15-18 ladda ner pdf (2000 KB) Part 6, 13(1): 19-22 ladda ner pdf (2000 KB) Part 7, 14(1): 44-49 ladda ner pdf (2600 KB) Part 8, 14(2): 31-35 ladda ner pdf (2700 KB) Part 9, 17(1): 21-26 ladda ner pdf (2700 KB)


Broström D. & K. Soop, 2000-2001: Några intressanta lokaler i Ovansiljan I, II, III — Jordstjärnan 21(2): 3-10; 22(2): 12-18 and 25(1): 21-31.

Summary:
A few interesting localities in the Upper Siljan Lake district, Dalecarlia, Sweden.
The following species are described and illustrated by colour photos.

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Part II: ladda ner pdf (2700 KB) Part III: ladda ner pdf (3800 KB)


Lindström, H. & Soop K., 1995: Cortinarius testaceofolius spec. nova, a common Telamonia of the taiga — Karstenia 35: 91-95.

Summary:
Cortinarius testaceofolius Lindstr. & Soop, sp. nov. is described from Sweden and Norway. The species is characterised principally by its conspicuous brick-red gills, and is associated with Picea abies, more rarely with Pinus sylvestris. It seems to be one of the most common members of subg. Telamonia in oligotrophic spruce forests of the northern taiga region.


Soop K., Strid Å., Toresson H.-G., 1995: Pleurotus calyptratus — slöjmusslingen — återfunnen i Sverige efter ett och ett halvt sekel — Jordstjärnan 16(3): 55-61.

Summary:
Pleurotus calyptratus found again in Sweden after one and a half century. Pleurotus calyptratus (Lindbl.) Sacc., one of three members of the genus with a veil and apparently very rare, was encountered in Sweden in 1995 (with colour photo).


Brandrud T.E, Knutsson T., Soop K., 1997: En fantastisk säsong på Öland. Rapport med tonvikt på Phlegmacium (spindelskivlingar) — Jordstjärnan 17(3): 27-42.
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Summary:
A fantastic season on Öland, SE Sweden — A report with emphasis on Phlegmacium (Cortinarius).
Due to an unusually rainy year 1996 in SE Sweden, some dry localities on the calcareous island of Öland exhibited an unprecedented development of Cortinarius subgenus Phlegmacium. This paper presents two of the most favourable localities and a species list. The list includes five taxa not previously published from Sweden and ten that have merely 1 to 4 known localities in the country, occurring mainly on Öland. Of the taxa, 24 are rare to very rare in Sweden and most of these are listed in the Swedish red list of fungi. The localities are characterised by dry, extensively grazed areas displaying a mosaic of shrubs, mainly of Corylus and Juniperus. Between the shrubs are dry grassland communities (Avenetum) with dominating grasses and herbs such as Avena pratensis, Helianthemum nummularium, and Thymus serpyllifolia. The localities are exposed to a very warm local climate and have a long continuity of mixed Corylus/Juniperus populations with sparse occurrences of Quercus, Betula, etc. These factors, along with the extremely calcareous soil, are responsible for the very diverse fungus flora, including more southerly species as well as species that normally occur in deciduous woodlands. The habitats are extremely vulnerable and under serious threat. Local authorities are urged to protect the remains of this type of eco-system and land-use. The vegetation, including the fungi, should be investigated further. During just a few visits 1995-1996, the best locality produced 150 recorded species of Basidiomycetes, of which 28 are in the Swedish red list of rare and threatened fungi. Most probably the type of fungal community found on the sites is unique in Europe. Further, a most valuable subject for investigation would be to study the association between mycorrhizal species and various species of shrubs and herbs, including Helianthemum spp. Taxa illustrated by colour photographs:


Soop K., 2012: Slöjfällskivling (Lepiota cortinarius) funnen i Sverige — Sv. Mykol. Tids. 33:2.
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Summary:
Lepiota cortinarius found in Sweden. The first confirmed records of Lepiota cortinarius J.E. Lange from Sweden are reported. They were collected in Picea forests in the boreal zone, indicating that the species is perhaps less continental than formerly believed.


Soop K., 1993: Schematisk nyckel till Cortinarius i Nord- och Mellansverige — Jordstjärnan 14(1): 4-28.


Matli F., Soop K., Toresson H.-G., 2006: Intressanta fynd av kamskivlingar — Svensk Mykologisk Tidskrift 27(2): 11-16.
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Sammanfattning
Vi rapporterar fynd av tre flugsvampar i undersläktet Amanitopsis från Mellan- och Sydsverige: Amanita islandica Melot, A. cf. ochraceomaculata Néville & al., och A. cf. pachycolea Stunz. Endast den första arten är slutgiltigt bestämd, medan de övriga är approximativa. Fynden illustreras med färgfoton.

Summary:
Interesting finds of Amanitopsis species. Three findings of Amanita in subgenus Amanitopsis are reported from South and Central Sweden : Amanita islandica Melot, A. cf. ochraceomaculata Néville & al., and A. cf. pachycolea Stunz. Only the first-mentioned species is fully idenitifed, while the other two are approximate. The collections are illustrated by colour photos.


Garnica S, Schön ME, Abarenkov K, Riess K, Liimatainen K, Niskanen T, Balint D, Soop K, Frøslev T, Jeppesen T, Peintner U, Kuhnert-Finkernagel R, Brandrud TE, Saar G, Oertel B, Ammirati J. 2016. Determining threshold values for barcoding fungi: lessons from Cortinarius (Basidiomycota), a highly diverse and widespread ectomycorrhizal genus. FEMS Microbiology Ecology 92(4).

Abstract:
Different distance-based threshold selection approaches were used to assess and compare use of the internal transcribed spacer (ITS) region to distinguish among 901 Cortinarius species represented by >3000 collections. Sources of error associated with genetic markers and selection approaches were explored and evaluated using MOTUs from genus and lineage based-alignments. Our study indicates that 1%Ð2% more species can be distinguished by using the full-length ITS barcode as compared to either the ITS1 or ITS2 regions alone. Optimal threshold values for different picking approaches and genetic marker lengths inferred from a subset of species containing major lineages ranged from 97.0% to 99.5% sequence.


Dima B, Lindström H, Liimatainen K, Olson , Soop K, Kytövuori I, Dahlberg A. Niskanen T. 2016. Typification of Friesian names in Cortinarius sections Anomali, Spilomei, and Bolares, and description of two new species from northern Europe. Mycol. Progress 15(9): 903-919.

Abstract:
Abstract Seven commonly used Friesian names of Cortinarius sect. Anomali, Spilomei, and Bolares are discussed. Neotypes for C. anomalus, C. spilomeus, and C. bolaris, as well as epitypes for C. albocyaneus, C. caninus, and C. tabularis are proposed. The distinction of these six species is supported by both morphological and rDNA internal transcribed spacer (ITS) data, whereas C. azureus, as currently typified, is considered as a synonym of C. anomalus, but could also be considered as nomen dubium. Barcodes (ITS) and morphological descriptions based on materials from a broad geographical area in Europe are provided for each species. Furthermore, two new species, C. jonimitchelliae and C. pastoralis, from northern Europe are described. The latter is proposed to accommodate C. anomalus subsp. campestris, which has been described from treeless habitats with Helianthemum. Our study reveals that the subsp. campestris is not an intraspecific variety of C. anomalus, but an own species close to C. caninus, and has a much wider distribution and host range than earlier thought. An identification key for the species/sections included here is provided.


Brandrud TE, Schmidt-Stohn G, Liimatainen K, Niskanen T, Frøslev TG, Soop K, Bojantchev D, Kytövuori I, Jeppesen TS, Bellù F, Saar G, Oertel B, Ali T, Thines M, Dima B. 2018. Cortinarius sect. Riederi: taxonomy and phylogeny of the new section with European and North American distribution. Mycol. Progress 17(12): 1323-1354.

Abstract:
Abstract Seven commonly used Friesian names of Cortinarius sect. Anomali, Spilomei, and Bolares are discussed. Neotypes for C. anomalus, C. spilomeus, and C. bolaris, as well as epitypes for C. albocyaneus, C. caninus, and C. tabularis are proposed. The distinction of these six species is supported by both morphological and rDNA internal transcribed spacer (ITS) data, whereas C. azureus, as currently typified, is considered as a synonym of C. anomalus, but could also be considered as nomen dubium. Barcodes (ITS) and morphological descriptions based on materials from a broad geographical area in Europe are provided for each species. Furthermore, two new species, C. jonimitchelliae and C. pastoralis, from northern Europe are described. The latter is proposed to accommodate C. anomalus subsp. campestris, which has been described from treeless habitats with Helianthemum. Our study reveals that the subsp. campestris is not an intraspecific variety of C. anomalus, but an own species close to C. caninus, and has a much wider distribution and host range than earlier thought. An identification key for the species/sections included here is provided.


Voto P, Soop K. 2018. Pseudobaeospora aciculifera, a new species from New Zealand. RMR, Boll. Amer 103: 23-26.

Abstract:
A new species of Pseudobaeospora collected in New Zealand is described; it is characterized, among other parameters, by the presence of pleurocystidioid elements, a micro morphological feature never known before to this genus. Descriptions and photos of macro- and microscopic features are presented.


Soop K, Schmidt-Stohn G, Dima B, Saar G, Brandrud TE. 2019. Cortinarius subgenus Phlegmacium section Aureocistophili in Europe. Journal des JEC 21: 33-54.

Abstract:
The taxonomy and phylogeny of sect. Aureocistophili (Cortinarius, subgen. Phlegmacium) are presented. The section is well circumscribed by molecular data and morphological features. We identify nine species in the section, two of which are undescribed. The study is based largely on data from our own collections; however, in three cases (C. aureocistophilus, C. kytoevuorii, and C. inusitatus) descriptions follow the protologues. In the study we generated nineteen new sequences, which serve as the base for a phylogram covering the section and other relevant portions of the subgenus. Species of section Aureocistophili are widespread all over Europe, from Fennoscandia to the Mediterranean. They grow on soils of varied acidity and with various coniferous and deciduous trees. Although more material is needed for some species, we publish here a provisional key of the section.


Soop K, Dima B, Cooper JA, Duckchul P, Oertel B. 2019. A phylogenetic approach to a global supraspecific taxonomy of Cortinarius (Agaricales) with an emphasis on the southern mycota. Persoonia 42: 261-290.

Abstract:
A section-based taxonomy of Cortinarius, covering large parts of the temperate North and South Hemispheres, is presented. Thirty-seven previously described sections are reviewed, while another forty-two sections are proposed as new or as new combinations. Twenty additional clades are recovered but not formally described. Furthermore, six new or combined species names are introduced, and one species is neotypified. The structure is supported by morphological characters and molecular evidence, based on two (nrITS and nrLSU) and four (nrITS, nrLSU, rpb1 and rpb2) loci datasets and analysed by Maximum Likelihood methods (PhyML, RAxML). Altogether 789 Cortinarius samples were included in the study.


Dima B, Semwal VK. Soop K, Brandrud TE, Wei TZ, Xie ML, Yao YJ. 2020. Nomenclatorial novelties. Index Fungorum 1277.
The following species in Cortinarius section Rubucunduli are described:


Schmidt-Stohn G, Saar G, Soop K, Brandrud TE, Dima B. 2020. Cortinarius subgenus Phlegmacium subsection Varii in Europe. Journal des JEC 22: 27-49.

Abstract:
The taxonomy and phylogeny of the new subsect. Varii (Cortinarius, subgen. Phlegmacium) are presented. The subsection is well circumscribed by molecular data and to some degree also on morphological features. We identify seven species in the subsection, six of them occurring in Europe. The study is mainly based on data from our own collections except for C. variiformis s. str. and C. variosimilis, whose descriptions follow the protologues, too. In the study we generated eighteen new sequences, which serve as the base for a phylogram covering the subsection. Species of subsect. Varii are characterised by their strict mycorrhizal associations, at least in Europe, with a wide variety of mycorrhizal hosts apparently mainly or exclusively growing on basic soils. Although more material is needed for some species, we publish here a provisional key of the subsection.


Dima B, Soop K. 2020. The "xenosmatoid" group of Cortinarius (Agaricales) in New Zealand. Phytotaxa 438 (4): 223-236.

Abstract:
Cortinarius section Xenosmatae, originally based on solely morphological characters, was subsequently shown to contain phylogenetically distantly related species. The type species C. xenosma is a singleton, and this study aims to revise the other members of the section using combined molecular (nrDNA ITS and LSU) and morphological data. Based on phylogenetic analyses using RAxML, PhyML and Bayesian Inference and additional morphological features one new species (C. paraxenosma) and one new section (sect. Olorinati) are proposed. Furthermore sect. Carbonelli is extended and emended to include two former members of sect. Xenosmatae. A key to the species in New Zealand with xenosmatoid morphology is provided.


Soop K. 2020. En intressant grupp av spindlingar (Cortinarius, Agaricales) inom undersläktet Phlegmacium. Svensk Mykologisk Tidskrift 41 (2): 37-45.

Abstract:
Cortinarius section Phlegmacioides in Sweden. An interesting group of Cortinarius section Phlegmacioides is reviewed in the light of Swedish findings and recent molecular studies. A number of rare species in the section are described: C. sobrius, C. balteaticlavatus, C. balteatibulbosus, and forms of the more common C. pseudon¾vosus and C. balteatocumatilis are briefly discussed. A new species, C. aff. decumbens, possessing a unique macromorphology in the section, is described in the hope that it may trigger additional findings. A phylogram of the section and colour plates of several phenotypic forms are presented. (In Swedish).


Soop K. 2022. Ten genera to replace Cortinarius. translated from Journal des JEC.

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Soop K. 2022. Dix genres pour remplacer Cortinarius. Journal des JEC 24: 22-24.

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Nilsen AR, Wang XY, Soop K, Cooper JA, Ridley GS, Wallace M, Summerfield TC, Brown CM. 2020. Purple haze: Cryptic purple sequestrate Cortinarius in New Zealand. Mycologia 112: 588-605.

Abstract:
Cortinarius is a species-rich ectomycorrhizal genus containing taxa that exhibit agaricoid or sequestrate basidiome morphologies. In New Zealand, one of the most recognizable and common Cortinarius species is the purple sequestrate fungus, C. porphyroideus. We used genome skimming of the almost 100-y-old type specimen from C. porphyroideus to obtain the nuc rDNA internal transcribed spacer region ITS1-5.8S-ITS2 (ITS barcode) and partial nuc rDNA 28S (28S) sequences. The phylogenetic position of C. porphyroideus was established, and we found that it represents a rarely collected species. Purple sequestrate Cortinarius comprise multiple cryptic species in several lineages. We describe four new species of Cortinarius with strong morphological similarity to C. porphyroideus: Cortinarius diaphorus, C. minorisporus, C. purpureocapitatus, and C. violaceocystidiatus. Based on molecular evidence, Thaxterogaster viola is recognized as Cortinarius violaceovolvatus var. viola. These species are associated with Nothofagus (southern beech) and have very similar morphology to C. porphyroideus but are all phylogenetically distinct based on molecular data.